A once-in-a-100,000-years global heatwave melts the Antarctic ice, awakening DAI-KAIJU from its long sleep! A great battle between DAI-KAIJU, which is over 100 meters long, and a giant girl begins in Shinjuku.
The special opens with Tachibana Tōbei taking some children to a Kamen Rider roadshow. Just as he's reminiscing about all the heroic modified humans he's lived alongside- Ichigō, Nigō, V3, Riderman, X, Amazon, Stronger and Tackle- the first seven Riders gradually show up to greet him in their human guises, unrecognised by the crowds. When it's revealed that the kaijin onstage are 'real', not actors, the Riders transform to save the crowd and Rider actors, uniting their power to defeat the Delzer Army's true leader, Great General Darkness in his hideout beneath the stadium.
Kei Watanabe is a fifth grade elementary school student. He has inherited the hunter's blood to hunt yōkai. To save the world from being destroyed, Kei Watanabe challenges a war against the yōkai.
Recognizing the destructive power of its captive giant Ape, the military makes its own battle-ready A.I., Mecha Ape. But its first practical test goes horribly wrong, leaving the military no choice but to release the imprisoned giant ape to stop the colossal robot before it destroys downtown Chicago.
JSDF pilot Akane has a fateful encounter when a new Godzilla emerges in Tateyama. As a countermeasure, a cyborg named Kiryu is constructed from the remains of the original. The machine is discovered to harbor the restless soul of the original monster as Akane must learn to find value in her own life as well.
The spirit of Kiba the Dark Makai Knight influenced everything since the beginning. Though he called himself a Makai Knight, Kiba was no different to a Horror. Kiba's spirit resides inside what was once a grey colored Makai Armor that's filled with the dark knowledge of Makai Power. As Barago absorbed Horror after Horror, Kiba eventually deformed into an organically pure black armor with a cape. Kiba assumed full control over his host upon Meisha's defeat by Kouga.
Yapool has returned after being injured in Ultraman Ginga S. He masterminds several attacks on Shou's homeworld, the Victorian kingdom in an attempt to get to the Victorium Core. Ultraman Hikari appears and grants Shou the ability to transform into Knight Mode with the Knight Timbre. Now free from their Spark Doll forms, some of the Showa Ultras join Victory in his fight.
Zero finds himself trapped within a universal gap and faced against an impostor Ace. Following the battle, Darklops Zero was restrained once more by Herodia, remembering a figure named "Kaiser Belial" before deactivating. Rei was restrained after his failed attempt to intrude the alien base but was rescued by none other than his alternate self. Unlike the prime Rei, alternate Reimon was never able to control his powers, forced into the state of Reionic Burst until the original Rei punched Reimon, thus deactivating its Reionic Burst mode. Rei revealed that he lost his Rayblood instinct after Hyuga threw Seven's Eye Slugger, creating a scar on his chest which symbolizes the former's friendship with ZAP SPACY members. While searching for Rei, Hyuga instead finds his Neo Battle Nizer and an entrance to the alien base made by Litra. Although the fake Ace was destroyed, Zero try to find an escaped but was halted by Seven and Ultraman impostors.
Tomoya leisurely sketches the Ultra Monsters in his sketchbook but an unknown forces suddenly radiated the book. Misuzu, Kenta and Chigusa discover several Spark Dolls, which were thought to be sent into outer space but somehow returned to Earth. Hikaru soon arrives, and soon the group are transported into a virtual room by Tomoya's Gunpad. The room itself has several Spark Dolls in its collection. Tomoya introduces them to the Livepad, a tablet which allows its user to use UltLive simulations by simply scanning the Spark Doll via the tip of its pointer pen. The group begins experimenting with the new technology.
After the previous Godzilla attack, a miniature arms race ensues to collect his cells. Concerned over Godzilla's possible return, the Japanese government uses the cells to create a new bio-weapon, ANEB (Anti-Nuclear Energy Bacteria). They seeks the aid of geneticist Genshiro Shiragami, who's experiments result in a new mutation.
During an assignment, foreign correspondent Steve Martin spends a layover in Tokyo and is caught amid the rampage of an unstoppable prehistoric monster the Japanese call 'Godzilla'. The only hope for both Japan and the world lies on a secret weapon, which may prove more destructive than the monster itself.
Astronauts Glenn and Fuji investigate Planet X and encounter mysterious aliens known as the Xiliens, who ask Earth's people to help save their world from "Monster Zero". In exchange for borrowing Godzilla and Rodan, the Xiliens offer a cure for cancer. As Glenn investigates, he develops a romance with Miss Namikawa and uncovers the Xilien's true intentions.
The Go-Busters must protect the Tokyo Enetower from the evil Vaglass organization. However Enter's new creation Steamloid causes the Buddy Roids to rust, making them unable to fight or be used in the Buster Machines.
Mining engineer Shigeru investigates the disappearance and death of his fellow coworkers when prehistoric nymphs are discovered emerging from the mines. After an attack on the local village, Shigeru heads deeper into the mines only to make a more horrifying discovery in the form a prehistoric flying creature. Soon a second monster appears as the two converge in Fukuoka.
A submarine expedition to salvage the remains of Mechagodzilla is thwarted by a massive dinosaur named Titanosaurus. An Interpol investigation leads biologist Ichinose to uncover the work of Dr. Mafune and his mysterious daughter Katsura. Aligned with the Black Hole Aliens, Katsura's life becomes entwined with the resurrected machine.
Manga artist Gengo Odaka lands a job with the World Children's Land amusement park only to become suspicious of the organization when a garbled message is discovered on tapes. As Gengo and his team investigate, Godzilla and Anguirus quickly decipher the message and begin their own plan of action.
Both TyrannoRanger and AbaRed are attacking the Kyoryugers because of mind control by Space War God Boldos. Boldos uses the feelings of revenge from enemies defeated by heroes who uses the power of dinosaurs, creating Neo-Grifforzer and Neo-Geildon, clones of past enemies as well as Enter and Escape,enemies of the Go-Busters. Villains spanning generations are united into a single evil force. The Kyoryugers, who are fighting an unseen evil, meet with the Go-Busters, and the "Greatest Trap ever seen on Earth" is sprung. What rises up against the true dinosaurs, and the death of your best friends? Now, the Kyoryugers must try and use the "True Brave".
Reporter Goro Maki stumbles upon scientists conducting weather experiments on Sollgel Island in the South Seas. He discovers the island is inhabited by giant mantis and a woman named Saeko who's been cast away since the death of her father. The pair soon find a helpless infant monster that Godzilla must adopt and learn to raise as one of his own.
Primatologist Davis Okoye shares an unshakable bond with George, the extraordinarily intelligent, silverback gorilla who has been in his care since birth. But a rogue genetic experiment gone awry mutates this gentle ape into a raging creature of enormous size. To make matters worse, it’s soon discovered there are other similarly altered animals. As these newly created alpha predators tear across North America, destroying everything in their path, Okoye teams with a discredited genetic engineer to secure an antidote, fighting his way through an ever-changing battlefield, not only to halt a global catastrophe but to save the fearsome creature that was once his friend.
The final Yodonna special. Yodonna, is now living in the human world after her revival, accidentally ends up working with a group of female bank robbers.